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College of Arts and Sciences

Faculty and Staff

Ryan Rykaczewski

Title: Assistant Professor
Department: Biological Sciences; Earth, Ocean and Environment
College of Arts and Sciences
Phone: 803-777-8159
Office: EWS, Room 612
Resources: Curriculum Vitae [pdf]
School of the Earth, Ocean and Environment
Department of Biological Sciences
My website
Ryan Rykaczewski


I am interested in exploring the responses of ecosystem and fisheries production to past and future climate variability and climate change. This research involves consideration of theory, observations, and models.  Currently, I investigate the biogeochemical implications of changes in physical climate that are particularly robust in the projections of atmosphere-ocean general circulation models (e.g., meridional shifts in zonal winds, increases in ocean stratification, and changes in phenology). In the past, changes in primary production, carbon export, the structure of food webs, and fisheries have often correlated with simple physical climate measures such as SST. However, these relationships rarely reflect an understanding of the actual mechanisms relating the environment to the physics.


  • Biogeochemistry & Geochemistry
  • Climate Change
  • Earth Systems
  • Marine Ecology and Evolution
  • Oceanography


Fisheries Oceanography; Observation of long-term changes in ocean biogeochemistry and climate; Large-scale changes in oceanic and atmospheric properties of the North Pacific; Remote forcing of coastal conditions; Ocean acidification and deoxygenation; Size-structured interactions in ocean ecosystems


García-Reyes, M, T Lamont, WJ Sydeman, BA Black, RR Rykaczewski, SA Thompson, SJ Bograd. In Press. A comparison of modes of upwelling-favorable wind variability in the Benguela and California current ecosystems. Journal of Marine Systems. doi:10.1016/j.jmarsys.2017.06.002. Link to Journal of Marine Systems.

Lynch, AL, RG Asch, WWL Cheung, CP Paukert, RR Rykaczewski, and WHH Sauer. 2017. Editorial for special issue: Impacts of climate change on marine and inland fishes and fisheries. Reviews in Fisheries Biology and Fisheries 27(2):293-296, doi:10.1007/s11160-017-9483-0. Link to RFBF.

*Brady, RX, MA Alexander, NS Lovenduski, and RR Rykaczewski. 2017. Emergent anthropogenic trends in California Current upwelling. Geophysical Research Letters 44, doi:10.1002/2017GL072945. Link to GRL.

Tommasi, D, CA Stock, AJ Hobday, R Methot, IC Kaplan, JP Eveson, K Holsman, TJ Miller, S Gaichas, M Gehlen, A Pershing, GA Vecchi, R Msadek, T Delworth, CM Eakin, MA Haltuch, R Sefarian, CM Spillman, JR Hartog, S Siedlecki, JF Samhouri, B Muhling, RG Asch, ML Pinsky, VS Saba, SB Kapnick, CF Gaitan, RR Rykaczewski, MA Alexander, Y Xue, KV Pegion, P Lynch, MR Payne, T Kristiansen, P Lehodey, and FE Werner. 2017. Managing living marine resources in a dynamic environment: the role of seasonal to decadal climate forecasts. Progress in Oceanography 152:15-49, doi:10.1016/j.pocean.2016.12.011. Link to Prog. Oceanogr.

Stock, CA, JG John, RR Rykaczewski, RG Asch, WWL Cheung, JP Dunne, KD Friedland, VWY Lam, JL Sarmiento, RA Watson. 2017. Reconciling fisheries catch and ocean productivity. Proceedings of the National Academy of Sciences of the United States of America 114:E1441-E1449, doi:10.1073/pnas.1610238114. Link to PNAS.

Checkley, DM Jr., RG Asch, and RR Rykaczewski. 2017. Climate, anchovy, and sardine. Annual Review of Marine Science 9:469-493, doi:10.1146/annurev-marine-122414-033819. Link to ARMS.

*Grieve, BD, EN Curchitser, and RR Rykaczewski. 2016. Range expansion of the invasive lionfish in the Northwest Atlantic with climate change. Marine Ecology Progress Series 546:225-237, doi:10.3354/meps11638. Link to MEPS.

Cheung, WWL, RG Asch, TL Frölicher, G Reygondeau, M Jones, ML Pinsky, KB Rodgers, RR Rykaczewski, JL Sarmiento, CA Stock, and JR Watson. 2016. Building confidence in projections of the responses of living marine resources to climate change. ICES Journal of Marine Science 73:1283-1296, doi: 10.1093/icesjms/fsv250. Link to ICES Journal of Marine Science.

García-Reyes, M, WJ Sydeman, DS Schoeman, RR Rykaczewski, BA Black, AJ Smit, and SJ Bograd. 2015. Under pressure: Climate change, upwelling and eastern boundary upwelling ecosystems. Frontiers in Marine Science 2:109, doi:10.3389/fmars.2015.00109. Link to Frontiers in Marine Science.

Fisher, JL, WT Peterson, and RR Rykaczewski. 2015. The impact of El Niño events on the pelagic food chain in the northern California Current. Global Change Biology 21:4401–4414, doi:10.1111/gcb.13054.  Link to GCB.

Rykaczewski, RR, JP Dunne, WJ Sydeman, M García-Reyes, BA Black, and SJ Bograd. 2015. Poleward displacement of coastal upwelling-favorable winds in the ocean's eastern boundary currents through the 21st century. Geophysical Research Letters 42:6424–6431, doi:10.1002/2015GL064694. Link to GRL.

Bakun, A, BA Black, SJ Bograd, M García-Reyes, AJ Miller, RR Rykaczewski, and WJ Sydeman. 2015. Anticipated effects of climate change on coastal upwelling ecosystems. Current Climate Change Reports 1:85-93, doi:10.1007/s40641-015-0008-4. Link to Current Climate Change Reports.

*Willis-Norton, E, EL Hazen, S Fossette, G Shillinger, RR Rykaczewski, DG Foley, JP Dunne, and SJ Bograd. 2015. Climate change impacts on leatherback turtle pelagic habitat in the southeast Pacific. Deep-Sea Research Part II Top Stud Oceanogr 113:260-267, doi:10.1016/j.dsr2.2013.12.019. Link to DSRII.

Black, BA, WJ Sydeman, DC Frank, D Griffin, DW Stahle, M García-Reyes, RR Rykaczewski, SJ Bograd, and WT Peterson. 2014. Six centuries of variability and extremes in a coupled marine-terrestrial ecosystem. Science 345(6203):1498-1502, doi:10.1126/science.1253209. Link to Science. Press Release.

Sydeman, WJ, M García-Reyes, DS Schoeman, RR Rykaczewski, SA Thompson, BA Black, SJ Bograd. 2014. Climate change and wind intensification in coastal upwelling ecosystems. Science 345(6192):77-80, doi:10.1126/science.1251635. Link to Science. Press Release. Los Angeles Times article.

McOwen, CJ, WWL Cheung, RR Rykaczewski, RA Watson, and LJ Wood. 2014. Is fisheries production within Large Marine Ecosystems determined by bottom-up or top-down forcing?  Fish and Fisheries 16:623–632, doi:10.1111/faf.12082. doi: 10.1111/faf.12082. 2014. Link to Fish & Fisheries.

García-Reyes, M, WJ Sydeman, BB Black, RR Rykaczewski, DS Schoeman, SA Thompson, and SJ Bograd. 2013. Relative influence of oceanic and terrestrial pressure systems in driving upwelling-favorable winds. Geophysical Research Letters 40:5311–5315, doi:10.1002/2013GL057729. Link to GRL.

Österblom, H, A Merrie, M Metian, WJ Boonstra, T Blenckner, JR Watson, RR Rykaczewski, Y Ota, JL Sarmiento, V Christensen, M Schlüter, S Birnbaum, BG Gustavsson, C Humborg, C Mörth, B Müller-Karulis, MT Tomczak, M Troel, C Folke. Modeling social-ecological scenarios in marine systems. 2013. BioScience 63:735-744, doi:10.1525/bio.2013.63.9.9. Link to BioScience. Press Release.

García-Reyes, M, WJ Sydeman, SA Thompson, BA Black, RR Rykaczewski, JA Thayer, and SJ Bograd. 2013. Integrated assessment of wind effects on Central California’s pelagic ecosystem. Ecosystems 16(5):722-735, doi: 10.1007/s10021-013-9643-6. Link to Ecosystems.

Beaulieu, C, SA Henson, JL Sarmiento, JP Dunne, SC Doney, RR Rykaczewski, and L Bopp. 2013. Factors challenging our ability to detect long-term trends in ocean chlorophyll. Biogeosciences 10:2711-2724, doi:10.5194/bg-10-2711-2013. Link to Biogeosciences.

Hazen, EL, E Jorgensen, RR Rykaczewski, SJ Bograd, DG Foley, ID Jonsen, SA Shaffer, JP Dunne, DP Costa, LB Crowder, and BA Block. 2013. Predicted habitat shifts of Pacific top predators in a changing climate. Nature Climate Change 3:234-238, doi:10.1038/nclimate1686. Link to Nature Climate Change. Press Release.

Sydeman, WJ, SA Thompson, JA Santora, JC Field, WT Peterson, RW Tanasichuk, HJ Freeland, SJ Bograd, and RR Rykaczewski. 2011. Does positioning of the North Pacific Current affect downstream ecosystem productivity? Geophysical Research Letters 38:L12606, doi:10.1029/2011GL047212. Link to GRL.

Rykaczewski, RR and JP Dunne. 2011. A measured look at ocean chlorophyll trends. Nature 472:E5-E6, doi:10.1038/nature09952. Link to Nature. New York Times "Dot Earth" blog.

Decima, M, MR Landry, and RR Rykaczewski. 2011. Broad-scale patterns in mesozooplankton biomass and grazing in the eastern equatorial Pacific. Deep-Sea Research Part II. 58:387-399, doi:10.1016/j.dsr2.2010.08.006. Link to DSRII.

Stock, CA, MA Alexander, NA Bond, K Brander, WWL Cheung, EN Curchitser, TL Delworth, JP Dunne, SM Griffies, MA Haltuch, JA Hare, AB Hollowed, P Lehodey, SA Levin, JS Link, KA Rose, RR Rykaczewski, JL Sarmiento, RJ Stouffer, FB Schwing, GA Vecchi, FE Werner. 2011. On the use of IPCC-class models to assess the impact of climate on living marine resources. Progress in Oceanography 88:1-27, doi:10.1016/j.pocean.2010.09.001. Link to Prog. Oceanogr.

Rykaczewski, RR and JP Dunne. 2010. Enhanced nutrient supply to the California Current Ecosystem with global warming and increased stratification in an earth system model. Geophysical Research Leters 37:L21606, doi:10.1029/2010GL045019. Link to GRL.

Rykaczewski, RR and DM Checkley. 2008. Influence of ocean winds on the pelagic ecosystem in upwelling regions. Proceedings of the National Academy of Sciences of the United States of America 105:1965-1970. Link to PNAS. Press Release.